Can a Great Dane ride comfortably in a Ford Fusion?

They're a bit small, but maybe if I put the seats down?

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Open up your window just enough for him to stick his head out.

Got you, E.H Amos
and Verulam

It is not safe or advisable to transport a dog who is not (at least) seat belted in. Can you do that, with the seats folded down? If not, and if no crate will fit in the vehicle, the dog will go F-L-Y-I-N-G, if you stop suddenly or are HIT by somebody else. There's also no safe way to leave the dog in the car if the weather is even slightly warm, if he can't be restrained and you can't open the windows.

What is the back of your car like if you lower the rear seats - too many have a step which won't leave enough space for a crate, which you really should have to transport any dog.

I'd not think this model of Ford, is going to be big enough.

Unless its the Estate Version of the Fusion, I can't see a Great Dane being very comfortable or safe in the car version of it. Jmo.

In my opinion no… It should be in a transport cage for its own benefit in a moving vehicle anyway